1 Office of the Director, National Institutes of Health, Bethesda, Maryland, United States of America.
2 Department of Plant and Microbial Biology, University of California, Berkeley, Berkeley, California, United States of America; Department of Molecular and Cell Biology, University of California, Berkeley, Berkeley, California, United States of America.
3 Department of Molecular and Cell Biology, University of California, Berkeley, Berkeley, California, United States of America.