1 Center for Retrovirus Research, The Ohio State University, Columbus, OH, USA.
2 Department of Veterinary Biosciences, The Ohio State University, Columbus, OH, USA.
3 Department of Microbial Infection and Immunity, The Ohio State University, Columbus, OH, USA.
4 Viruses and Emerging Pathogens Program, Infectious Diseases Institute, The Ohio State University, Columbus, OH, USA.
5 Food Animal Health Research Program, Ohio Agricultural Research and Development Center, CFAES, Department of Veterinary Preventive Medicine, The Ohio State University, Wooster, OH, USA.
6 Department of Microbiology, Perelman School of Medicine, University of Pennsylvania, Philadelphia, PA, USA.
7 Lineberger Comprehensive Cancer Center, Department of Microbiology and Immunology, University of North Carolina at Chapel Hill, Chapel Hill, NC, USA.