1 Biodiversity Research Center, Academia Sinica, Taipei, 115, Taiwan.
2 Institute of Ecology and Evolutionary Biology, College of Life Science, National Taiwan University, Taipei, 115, Taiwan.
3 Department of Ecology, Evolution and Environmental Biology and Center for Integrative Animal Behavior, Columbia University, New York, NY, 10027, USA.
4 Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu, 61004, People's Republic of China.
5 Department of Life Sciences, National Cheng Kung University, Tainan, 70101, Taiwan. chenic@mail.ncku.edu.tw.
6 Biodiversity Research Center, Academia Sinica, Taipei, 115, Taiwan. shensf@sinica.edu.tw.
7 Institute of Ecology and Evolutionary Biology, College of Life Science, National Taiwan University, Taipei, 115, Taiwan. shensf@sinica.edu.tw.