Abstract
Human Toll-like receptor (TLR) 3 recognizes double-stranded (ds) RNA and induces production of interferon (IFN)-beta independent of the adaptor molecules MyD88 and TIRAP. Thus, another adaptor must exist that preferentially mediates TLR3-dependent production of IFN-beta. We have identified an alternative adaptor, designated Toll-interleukin 1 receptor domain (TIR)-containing adaptor molecule (TICAM)-1, that can physically bind the TIR domain of TLR3 and activate the IFN-beta promoter in response to poly(I):poly(C). Thus, dsRNA-TLR3-dependent production of IFN-beta is mediated mainly by TICAM-1. This TICAM-1-dependent pathway may have a role in other TLR-IFN-beta pathways, which form part of the MyD88-independent cellular immune response.
Publication types
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Research Support, Non-U.S. Gov't
MeSH terms
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Adaptor Proteins, Signal Transducing
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Adaptor Proteins, Vesicular Transport / genetics
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Adaptor Proteins, Vesicular Transport / immunology*
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Adaptor Proteins, Vesicular Transport / metabolism*
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Amino Acid Sequence
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Animals
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Antigens, Differentiation / metabolism
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Binding Sites
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Cell Line
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Cloning, Molecular
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DNA-Binding Proteins / metabolism
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Drosophila Proteins*
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Female
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Gene Silencing
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Humans
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Immunity, Cellular
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Interferon Regulatory Factor-3
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Interferon-beta / biosynthesis*
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Male
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Membrane Glycoproteins / chemistry
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Membrane Glycoproteins / metabolism*
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Mice
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Molecular Sequence Data
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Myeloid Differentiation Factor 88
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NF-kappa B / metabolism
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Pregnancy
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Protein Structure, Tertiary
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RNA, Double-Stranded / metabolism
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Receptors, Cell Surface / chemistry
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Receptors, Cell Surface / metabolism*
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Receptors, Immunologic / metabolism
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Receptors, Interleukin-1 / metabolism
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Sequence Homology, Amino Acid
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Signal Transduction
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Tissue Distribution
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Toll-Like Receptor 3
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Toll-Like Receptors
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Transcription Factor AP-1 / metabolism
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Transcription Factors / metabolism
Substances
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Adaptor Proteins, Signal Transducing
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Adaptor Proteins, Vesicular Transport
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Antigens, Differentiation
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DNA-Binding Proteins
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Drosophila Proteins
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IRF3 protein, human
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Interferon Regulatory Factor-3
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Irf3 protein, mouse
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MYD88 protein, human
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Membrane Glycoproteins
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Myd88 protein, mouse
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Myeloid Differentiation Factor 88
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NF-kappa B
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RNA, Double-Stranded
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Receptors, Cell Surface
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Receptors, Immunologic
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Receptors, Interleukin-1
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TICAM-1 protein, mouse
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TICAM1 protein, human
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TIRAP protein, human
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TIRAP protein, mouse
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TLR3 protein, human
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Toll-Like Receptor 3
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Toll-Like Receptors
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Transcription Factor AP-1
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Transcription Factors
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Interferon-beta
Associated data
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GENBANK/AB086380
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GENBANK/AB091053