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Basal Tyrannosauroids From China and Evidence for Protofeathers in Tyrannosauroids

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Basal Tyrannosauroids From China and Evidence for Protofeathers in Tyrannosauroids

Xing Xu et al. Nature.

Abstract

Tyrannosauroids are one of the last and the most successful large-bodied predatory dinosaur groups, but their early history remains poorly understood. Here we report a new basal tyrannosauroid from the Early Cretaceous Yixian Formation of western Liaoning, China, which is small and gracile and has relatively long arms with three-fingered hands. The new taxon is the earliest known unquestionable tyrannosauroid found so far. It shows a mosaic of characters, including a derived cranial structure resembling that of derived tyrannosauroids and a primitive postcranial skeleton similar to basal coelurosaurians. One of the specimens also preserves a filamentous integumentary covering similar to that of other coelurosaurian theropods from western Liaoning. This provides the first direct fossil evidence that tyrannosauroids had protofeathers.

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