Testosterone stimulates Duox1 activity through GPRC6A in skin keratinocytes

J Biol Chem. 2014 Oct 17;289(42):28835-45. doi: 10.1074/jbc.M114.583450. Epub 2014 Aug 27.

Abstract

Testosterone is an endocrine hormone with functions in reproductive organs, anabolic events, and skin homeostasis. We report here that GPRC6A serves as a sensor and mediator of the rapid action of testosterone in epidermal keratinocytes. The silencing of GPRC6A inhibited testosterone-induced intracellular calcium ([Ca(2+)]i) mobilization and H2O2 generation. These results indicated that a testosterone-GPRC6A complex is required for activation of Gq protein, IP3 generation, and [Ca(2+)]i mobilization, leading to Duox1 activation. H2O2 generation by testosterone stimulated the apoptosis of keratinocytes through the activation of caspase-3. The application of testosterone into three-dimensional skin equivalents increased the apoptosis of keratinocytes between the granular and stratified corneum layers. These results support an understanding of the molecular mechanism of testosterone-dependent apoptosis in which testosterone stimulates H2O2 generation through the activation of Duox1.

Keywords: Apoptosis; Cell Signaling; NADPH Oxidase; Reactive Oxygen Species (ROS); Testosterone.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Apoptosis
  • Calcium / metabolism
  • Caspase 3 / metabolism
  • Cell Line
  • Dual Oxidases
  • Gene Expression Profiling
  • Gene Expression Regulation, Enzymologic*
  • Humans
  • Hydrogen Peroxide / metabolism
  • Keratinocytes / metabolism*
  • Membrane Potential, Mitochondrial
  • NADPH Oxidases / metabolism*
  • Oxidation-Reduction
  • Receptors, G-Protein-Coupled / metabolism*
  • Signal Transduction
  • Skin / cytology
  • Skin / drug effects
  • Skin / metabolism*
  • Testosterone / chemistry*

Substances

  • GPRC6A protein, human
  • Receptors, G-Protein-Coupled
  • Testosterone
  • Hydrogen Peroxide
  • Dual Oxidases
  • NADPH Oxidases
  • DUOX1 protein, human
  • CASP3 protein, human
  • Caspase 3
  • Calcium