1 School of Geography, Planning and Environmental Management, The University of Queensland, St Lucia Qld 4072, Australia; Australian Centre for Ecological Analysis and Synthesis, Terrestrial Ecosystem Research Network, Australia.
2 James Hutton Institute, Invergowrie Dundee DD2 5DA, Scotland, UK.
4 School of Plant Biology, University of Western Australia, Crawley, WA 6009, Australia.
5 School of Geography, Planning and Environmental Management, The University of Queensland, St Lucia Qld 4072, Australia; Terrestrial Ecosystem Research Network, Goddard Building, University of Queensland, Australia.