Megakaryocytic hyperplasia in myeloproliferative neoplasms is driven by disordered proliferative, apoptotic and epigenetic mechanisms

J Clin Pathol. 2016 Feb;69(2):155-63. doi: 10.1136/jclinpath-2015-203177. Epub 2015 Aug 19.

Abstract

Aims: Myeloproliferative neoplasms (MPN) are a heterogeneous group of clonal proliferative bone marrow diseases characterised by extensive megakaryocytic hyperplasia and morphological atypia. Despite knowledge of genomic defects, the pathobiological processes driving these megakaryocytic abnormalities in MPN remain poorly explained. We have explored the proliferative, apoptotic and epigenetic profiles of megakaryocytes in human MPN.

Methods: Immunohistochemical staining was performed on bone marrow trephine biopsies of 81 MPN (with and without JAK2(V617F) and CALR mutations) and 15 normal controls to assess the megakaryocytic expression of biomarkers associated with proliferation (Ki67), apoptosis (Bcl-XL, BNIP-3) and epigenetic regulation (EZH2, SUZ12).

Results: Myeloproliferative megakaryocytes showed significantly greater expression of proliferative Ki67 and anti-apoptotic Bcl-XL, reduced pro-apoptotic BNIP-3 and increased SUZ12 compared with controls. In essential thrombocythaemia, large-giant megakaryocytes with hyperlobated nuclei showed a trend towards a proliferative signature. In contrast, myelofibrotic megakaryocytes with condensed nuclear chromatin, and cases with CALR mutations, had significant reductions in pro-apoptotic BNIP-3.

Conclusions: Uncontrolled megakaryocytic expansion in MPN results from a combination of increased proliferation, attenuated apoptosis and defective epigenetic regulation with CALR mutations favouring apoptotic failure. The higher platelet counts reported to be seen in MPN with CALR mutations may be due to greater dysregulation of megakaryocyte apoptosis.

Keywords: APOPTOSIS; BONE MARROW; PROLIFERATION; fibrosis.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Apoptosis*
  • Biomarkers, Tumor / analysis
  • Biomarkers, Tumor / genetics
  • Biopsy
  • Bone Marrow Examination
  • Calreticulin / genetics
  • Case-Control Studies
  • Cell Proliferation*
  • DNA Mutational Analysis
  • Enhancer of Zeste Homolog 2 Protein
  • Epigenesis, Genetic*
  • Humans
  • Hyperplasia
  • Immunohistochemistry
  • Janus Kinase 2 / genetics
  • Ki-67 Antigen / analysis
  • Megakaryocytes / chemistry
  • Megakaryocytes / pathology*
  • Membrane Proteins / analysis
  • Mutation
  • Myeloproliferative Disorders / genetics
  • Myeloproliferative Disorders / metabolism
  • Myeloproliferative Disorders / pathology*
  • Neoplasm Proteins
  • Polycomb Repressive Complex 2 / analysis
  • Proto-Oncogene Proteins / analysis
  • Tissue Array Analysis
  • Transcription Factors
  • bcl-X Protein / analysis

Substances

  • BCL2L1 protein, human
  • BNIP3 protein, human
  • Biomarkers, Tumor
  • CALR protein, human
  • Calreticulin
  • Ki-67 Antigen
  • Membrane Proteins
  • Neoplasm Proteins
  • Proto-Oncogene Proteins
  • SUZ12 protein, human
  • Transcription Factors
  • bcl-X Protein
  • EZH2 protein, human
  • Enhancer of Zeste Homolog 2 Protein
  • Polycomb Repressive Complex 2
  • JAK2 protein, human
  • Janus Kinase 2