1 Department of Genetics, Evolution and Environment, University College London, London WC1E 6BT, UK.
2 Whitney Laboratory for Marine Bioscience, University of Florida, St Augustine, Florida 32080, USA, and in the Department of Neuroscience and at the McKnight Brain Institute, University of Florida, Gainesville.
3 Department of Biological Sciences, Auburn University, Auburn, Alabama 36849, USA.