1 King Abdullah University of Science and Technology (KAUST), Center for Desert Agriculture, Biological and Environmental Sciences and Engineering Division, Thuwal 23955-6900, Kingdom of Saudi Arabia.
3 Laboratoire de Recherche en Sciences Végétales, Université de Toulouse, CNRS, UPS, 24 chemin de Borderouge, Auzeville, BP 42617, 31326 Castanet-Tolosan, France.
4 King Abdullah University of Science and Technology (KAUST), Center for Desert Agriculture, Biological and Environmental Sciences and Engineering Division, Thuwal 23955-6900, Kingdom of Saudi Arabia. Electronic address: salim.babili@kaust.edu.sa.