1 Centre for Immunity, Infection and Evolution, University of Edinburgh, Edinburgh, UK; Usher Institute of Population Health Sciences and Informatics, University of Edinburgh, Edinburgh, UK.
2 Centre for Immunity, Infection and Evolution, University of Edinburgh, Edinburgh, UK.
3 Centre for Immunity, Infection and Evolution, University of Edinburgh, Edinburgh, UK; Regional Infectious Diseases Unit, Western General Hospital, Edinburgh, UK.
4 Usher Institute of Population Health Sciences and Informatics, University of Edinburgh, Edinburgh, UK.