Characterization of an oligosaccharide-pyrophosphodolichol pyrophosphatase activity in yeast

Biochem J. 1988 Oct 1;255(1):235-42.

Abstract

Use of Triton X-114 allowed us to develop a new method to separate hydrophilic oligosaccharidic material from hydrophobic oligosaccharide pyrophosphodolichols (oligosaccharide-PP-Dol). Taking advantage of this procedure we characterize, in yeast microsomal membranes, an enzymic activity that hydrolyses oligosaccharide-PP-Dol into oligosaccharidic material. H.p.l.c. analysis together with alkaline-phosphatase- and endo-N-acetyl-beta-D-glucosaminidase-susceptibility demonstrate that the oligosaccharidic released material is mainly composed of oligomannosides containing a chitobiose phosphate at the reducing end. The enzymic activity requires bivalent cations and is inhibited by pyrophosphate, NAD+ and bacitracin. As other, commercially available, pyrophosphatases have no action on lipid intermediates, the described pyrophosphatase activity appears to be the specific enzyme for oligosaccharide-PP-Dol. This enzymic splitting of the pyrophosphate bond might be the primary event in the catabolism of lipid intermediates.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Bacitracin / pharmacology
  • Cations, Divalent / pharmacology
  • Chromatography, Gel
  • Detergents / pharmacology
  • Glycosylation
  • Intracellular Membranes / enzymology
  • Microsomes / enzymology
  • NAD / pharmacology
  • Octoxynol
  • Oligosaccharides / metabolism
  • Phosphates / pharmacology
  • Polyethylene Glycols
  • Polyisoprenyl Phosphate Oligosaccharides / metabolism
  • Pyrophosphatases / antagonists & inhibitors
  • Pyrophosphatases / metabolism*
  • Saccharomyces cerevisiae / enzymology*

Substances

  • Cations, Divalent
  • Detergents
  • Oligosaccharides
  • Phosphates
  • Polyisoprenyl Phosphate Oligosaccharides
  • dolichyl diphosphate oligosaccharides
  • NAD
  • Bacitracin
  • Polyethylene Glycols
  • Octoxynol
  • Pyrophosphatases
  • oligosaccharide-diphosphodolichol pyrophosphatase