1 Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, 142 Xizhimenwai Street, Beijing, 100044, China; CAS Center for Excellence in Life and Paleoenvironment, Beijing, 100044, China; Department of Early Prehistory and Quaternary Ecology, Universität Tübingen, Tübingen, 72070, Germany. Electronic address: lifeng@ivpp.ac.cn.
2 Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, 142 Xizhimenwai Street, Beijing, 100044, China; CAS Center for Excellence in Life and Paleoenvironment, Beijing, 100044, China. Electronic address: chenfuyou@ivpp.ac.cn.
3 Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, 142 Xizhimenwai Street, Beijing, 100044, China; CAS Center for Excellence in Life and Paleoenvironment, Beijing, 100044, China.
4 Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, 142 Xizhimenwai Street, Beijing, 100044, China; CAS Center for Excellence in Life and Paleoenvironment, Beijing, 100044, China; University of Chinese Academy of Sciences, Beijing, 100049, China.