1 Behavioural Ecology, Institute of Ecology and Evolution, University of Bern, 3032 Hinterkappelen, Switzerland; sacha.engelhardt@iee.unibe.ch.
2 Behavioural Ecology and Physiological Group, Groningen Institute for Evolutionary Life Sciences, University of Groningen, 9700 CC Groningen, The Netherlands.
3 Department of Animal Sciences, Behavioural Ecology Group, Wageningen University & Research, 6700 AH Wageningen, The Netherlands.
4 Behavioural Ecology, Institute of Ecology and Evolution, University of Bern, 3032 Hinterkappelen, Switzerland.