1 School of Biological Sciences, Centre for Ecology, Evolution and Behaviour, Royal Holloway University of London, Egham, Surrey TW20 0EX, UK.
2 College of Animal Science and Technology, Northwest A&F University, Yangling, Shaanxi 712100, China.
3 Department of Biology and Ecology of Fishes, Leibniz-Institute of Freshwater Ecology and Inland Fisheries, Müggelseedamm 310, Berlin, D-12587, Germany.