Sodium channel TRPM4 and sodium/calcium exchangers (NCX) cooperate in the control of Ca2+-induced mucin secretion from goblet cells

J Biol Chem. 2019 Jan 18;294(3):816-826. doi: 10.1074/jbc.RA117.000848. Epub 2018 Nov 27.

Abstract

Regulated mucin secretion is essential for the formation of the mucus layer that protects the underlying epithelial cells from foreign particles. Alterations in the quantity or quality of secreted mucins are therefore detrimental to airway and colon physiology. Based on various biochemical assays in several human cell lines, we report here that Na+/Ca2+ exchanger 2 (NCX2) works in conjunction with transient receptor potential cation channel subfamily M member 4 (TRPM4), and perhaps TRPM5, Na+ channels to control Ca2+-mediated secretion of both mucin 2 (MUC2) and MUC5AC from HT29-18N2 colonic cancer cells. Differentiated normal bronchial epithelial (NHBE) cells and tracheal cells from patients with cystic fibrosis (CFT1-LC3) expressed only TRPM4 and all three isoforms of NCXs. Blocking the activity of TRPM4 or NCX proteins abrogated MUC5AC secretion from NHBE and CFT1-LC3 cells. Altogether, our findings reveal that NCX and TRPM4/TRPM5 are both required for mucin secretion. We therefore propose that these two proteins could be potential pharmacological targets to control mucus-related pathologies such as cystic fibrosis.

Keywords: MUC2; MUC5AC; TRPM4; TRPM5; cystic fibrosis; goblet cell; mucin; secretion; sodium-calcium exchange; transient receptor potential channels (TRP channels).

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Calcium / metabolism*
  • Cell Line
  • Cystic Fibrosis / genetics
  • Cystic Fibrosis / metabolism
  • Cystic Fibrosis / pathology
  • Goblet Cells / metabolism*
  • Goblet Cells / pathology
  • Humans
  • Mucin 5AC / genetics
  • Mucin 5AC / metabolism*
  • Mucin-2 / genetics
  • Mucin-2 / metabolism*
  • Sodium-Calcium Exchanger / genetics
  • Sodium-Calcium Exchanger / metabolism*
  • TRPM Cation Channels / genetics
  • TRPM Cation Channels / metabolism*

Substances

  • MUC2 protein, human
  • MUC5AC protein, human
  • Mucin 5AC
  • Mucin-2
  • Sodium-Calcium Exchanger
  • TRPM Cation Channels
  • TRPM4 protein, human
  • TRPM5 protein, human
  • Calcium