1 Interdisciplinary Computing and Complex Biosystems Research Group, School of Computing Science, Newcastle University, Newcastle upon Tyne, UK.
2 School of Computing, Newcastle University, Newcastle upon Tyne, UK.
3 Mathematics and Statistics, Newcastle University, Newcastle upon Tyne, UK.
4 National Institute for Health Research, Newcastle Biomedical Research Centre, Newcastle upon Tyne Hospitals NHS Foundation Trust and Newcastle University, Institute of Cellular Medicine (Musculoskeletal Research Group), Newcastle upon Tyne, UK.
5 Newcastle University, Institute of Cellular Medicine, Musculoskeletal Research Group, Newcastle University and Newcastle upon Tyne Hospitals NHS Foundation Trust, Newcastle upon Tyne, UK.
6 School of Natural and Environmental Sciences, Newcastle University, Newcastle upon Tyne, UK.