1 Centre for Genomic and Experimental Medicine, Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh, EH4 2XU, UK. riccardo.marioni@ed.ac.uk.
2 Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, EH8 9JZ, UK. riccardo.marioni@ed.ac.uk.
3 Centre for Genomic and Experimental Medicine, Institute of Genetics and Molecular Medicine, University of Edinburgh, Edinburgh, EH4 2XU, UK.
4 Centre for Cognitive Ageing and Cognitive Epidemiology, University of Edinburgh, Edinburgh, EH8 9JZ, UK.
5 Institute for Molecular Bioscience, University of Queensland, Brisbane, QLD, 4072, Australia.
6 Social, Genetic and Developmental Psychiatry Centre, Institute of Psychiatry, Psychology & Neuroscience, King's College London, London, SE5 8AF, UK.
7 Department of Psychology, University of Edinburgh, Edinburgh, EH8 9JZ, UK.
8 Centre for Dementia Prevention, Centre for Clinical Brain Sciences, University of Edinburgh, Edinburgh, EH8 9YL, UK.
9 MRC Lifecourse Epidemiology Unit, University of Southampton, Southampton, SO16 6YD, UK.
10 Alzheimer Scotland Dementia Research Centre, University of Edinburgh, Edinburgh, EH8 9JZ, UK.
11 Departments of Neuroscience, Neurology and Genetics and Genomic Sciences, Ronald M. Loeb Center for Alzheimer's disease, Icahn School of Medicine at Mount Sinai, New York, NY, 10029-5674, USA.
12 Queensland Brain Institute, University of Queensland, Brisbane, QLD, 4072, Australia.