Abstract
Novel COVID-19 therapeutics are urgently needed. We generated a phage-displayed human antibody VH domain library from which we identified a high-affinity VH binder ab8. Bivalent VH, VH-Fc ab8, bound with high avidity to membrane-associated S glycoprotein and to mutants found in patients. It potently neutralized mouse-adapted SARS-CoV-2 in wild-type mice at a dose as low as 2 mg/kg and exhibited high prophylactic and therapeutic efficacy in a hamster model of SARS-CoV-2 infection, possibly enhanced by its relatively small size. Electron microscopy combined with scanning mutagenesis identified ab8 interactions with all three S protomers and showed how ab8 neutralized the virus by directly interfering with ACE2 binding. VH-Fc ab8 did not aggregate and did not bind to 5,300 human membrane-associated proteins. The potent neutralization activity of VH-Fc ab8 combined with good developability properties and cross-reactivity to SARS-CoV-2 mutants provide a strong rationale for its evaluation as a COVID-19 therapeutic.
Keywords:
SARS-CoV-2; electron microscopy; human V(H) antibody domain; mouse and hamster models; virus neutralization.
Copyright © 2020 Elsevier Inc. All rights reserved.
Publication types
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Research Support, N.I.H., Extramural
MeSH terms
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Angiotensin-Converting Enzyme 2
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Animals
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Antibodies, Neutralizing / immunology
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Antibodies, Neutralizing / ultrastructure
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Antibodies, Viral / administration & dosage
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Antibodies, Viral / chemistry
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Antibodies, Viral / immunology
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Antibodies, Viral / ultrastructure
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Antibody Affinity
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COVID-19
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COVID-19 Drug Treatment
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Coronavirus Infections / drug therapy*
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Cricetinae
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Female
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Humans
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Immunoglobulin Fc Fragments / immunology
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Immunoglobulin Heavy Chains / administration & dosage*
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Immunoglobulin Heavy Chains / chemistry
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Immunoglobulin Heavy Chains / immunology
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Immunoglobulin Heavy Chains / ultrastructure
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Immunoglobulin Variable Region / administration & dosage*
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Immunoglobulin Variable Region / chemistry
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Immunoglobulin Variable Region / immunology
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Immunoglobulin Variable Region / ultrastructure
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Mice
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Mice, Inbred BALB C
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Mutation
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Pandemics
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Peptide Library*
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Peptidyl-Dipeptidase A / metabolism
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Pneumonia, Viral / drug therapy*
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Protein Domains
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Spike Glycoprotein, Coronavirus / chemistry
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Spike Glycoprotein, Coronavirus / genetics
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Spike Glycoprotein, Coronavirus / metabolism
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Spike Glycoprotein, Coronavirus / ultrastructure
Substances
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Antibodies, Neutralizing
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Antibodies, Viral
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Immunoglobulin Fc Fragments
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Immunoglobulin Heavy Chains
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Immunoglobulin Variable Region
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Peptide Library
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Spike Glycoprotein, Coronavirus
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spike protein, SARS-CoV-2
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Peptidyl-Dipeptidase A
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ACE2 protein, human
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Ace2 protein, mouse
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Angiotensin-Converting Enzyme 2