1 Department of Engineering Science, University of Oxford, Parks Road, Oxford, OX1 3PJ, UK.
2 State Key Laboratory of Microbial Resources, Institute of Microbiology, Chinese Academy of Sciences, Beijing, China.
3 College of Engineering, Peking University, Beijing, 100871, China.
4 State Key Laboratory of Microbial Technology, Shandong University, Qingdao, 266237, China.
5 State Key Laboratory of Microbial Metabolisms, School of Sciences and Biotechnology; School of Oceanography, Shanghai JiaoTong University, Minhang, Shanghai, 200240, China.