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. 2021 Feb 4;16(2):e0245700.
doi: 10.1371/journal.pone.0245700. eCollection 2021.

Short-term occupations at high elevation during the Middle Paleolithic at Kalavan 2 (Republic of Armenia)

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Free PMC article

Short-term occupations at high elevation during the Middle Paleolithic at Kalavan 2 (Republic of Armenia)

Ariel Malinsky-Buller et al. PLoS One. .
Free PMC article

Abstract

The Armenian highlands encompasses rugged and environmentally diverse landscapes and is characterized by a mosaic of distinct ecological niches and large temperature gradients. Strong seasonal fluctuations in resource availability along topographic gradients likely prompted Pleistocene hominin groups to adapt by adjusting their mobility strategies. However, the role that elevated landscapes played in hunter-gatherer settlement systems during the Late Pleistocene (Middle Palaeolithic [MP]) remains poorly understood. At 1640 m above sea level, the MP site of Kalavan 2 (Armenia) is ideally positioned for testing hypotheses involving elevation-dependent seasonal mobility and subsistence strategies. Renewed excavations at Kalavan 2 exposed three main occupation horizons and ten additional low densities lithic and faunal assemblages. The results provide a new chronological, stratigraphical, and paleoenvironmental framework for hominin behaviors between ca. 60 to 45 ka. The evidence presented suggests that the stratified occupations at Kalavan 2 locale were repeated ephemerally most likely related to hunting in a high-elevation within the mountainous steppe landscape.

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Conflict of interest statement

The authors have declared that no competing interests exist.

Figures

Fig 1
Fig 1. Location map of Kalavan 2 within southern Caucasus and north-eastern Armenian highlands showing main Lower, Middle, and Upper Paleolithic sites.
Squares = open-air sites, triangles = cave sites.
Fig 2
Fig 2. Geological map of Kalavan area showing the larger watershed of the Barepat and in blue line the Dani watershed.
Fig 3
Fig 3. Plateau topography and location of trenches at Kalavan 2.
Fig 4
Fig 4
A: Section drawing of the Kalavan 2, Trench 1 north section including sample locations; B: Section drawing of the Kalavan 2, Trench 1 south section including sample locations.
Fig 5
Fig 5. Kalavan 2, Trench 2 east and south sections with sample locations.
Fig 6
Fig 6. Kalavan 2, Trench 3 south section with sample locations.
Fig 7
Fig 7. Kalavan 2, Trench 4 north section with sample locations.
Fig 8
Fig 8. Obsidian lithic artifacts.
1, 4–5. Shaping flakes that removed the tips of convergent retouched points; 2–3. Shaping flakes; 6–7. Retouched points. (Illustration: P. Glauberman).
Fig 9
Fig 9
1. Non-obsidian lithic artifacts: blade made on welded tuff; 2. Cortical flake on welded tuff; 3. Levallois point made on basalt; 4. Core on split pebble made on metamorphic material. (Illustration: P. Glauberman).
Fig 10
Fig 10
Fragment of a retouched blade (upper left) and end-scaper (upper right) with striations and polish associated with hard animal material. A retouched point (bottom left, Fig 8:6) and a shaping flake of a retouched point (bottom right, Fig 8:1) showing striations and polish diagnostic of woodworking activities.
Fig 11
Fig 11
Limestone pitted anvil showing evidence of a) impact marks and b) surface crushing. Bottom: c) Limestone slab with abraded surface and percussion impact feature; d) Limestone pitted anvil with several impact marks and multidirectional flake scars resulting from percussive actions (arrows indicate the orientation of the flakes removal); e) The archeological context of the pitted anvils (red circle indicating pitted anvil; yellow circles indicating bone fragments).
Fig 12
Fig 12. Locations of obsidian sources relative to Kalavan 2 (SRTM3 digital elevation data).
Presence or absence of the obsidians at Kalavan 2 is represented by each source’s circular symbol. The empty circles correspond to obsidian sources that are not currently reflected in the Kalavan 2 assemblage.
Fig 13
Fig 13. Obsidian sources and the corresponding source areas identified at Kalavan 2 by trench.
Fig 14
Fig 14. Obsidian source areas identified in all trenches by lithic class.
Fig 15
Fig 15. Kalavan 2 cervid (red deer sized) metacarpus with multiple cut marks (Id: N50-44).
Fig 16
Fig 16. Kalavan 2: Large bovid femur diaphysis with impact notch (Id: A77-1).
Fig 17
Fig 17
Binary plots of artifact density and retouch frequency for sedimentary units at Barozh 12 (open circles) and Kalavan 2 (filled circles), with obsidian artifacts (A) and non-obsidian artifacts at Kalavan 2 (B). Data exclude artifacts < 2 cm in maximal dimensions to avoid differences among sedimentary units in assemblage preservation and winnowing of small artifacts due to post-depositional processes.

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Grants and funding

For the essential financial and similar support, we gratefully thank the funding agencies of The Gerda Henkel Stiftung grant n. AZ 10_V_17 and n. AZ 23/F/19, the Leakey Foundation, and the Gfoeller Renaissance Foundation of USA. RT, AC, and SB would like to acknowledge support from the Leverhulme Trust-funded Palaeolithic Archaeology, Geochronology, and Environments of the Southern Caucasus (PAGES) project, and to thank Dr. C. Hayward at the University of Edinburgh for assistance with the microprobe analyses. VO and OB were supported by the ECCOREV Research Federation (Aix-Marseille Université, CNRS) and Labex OT-Med (n° ANR- 11-LABX-0061) funded by the French Government “Investissements d’Avenir” programme of the French National Research Agency (ANR) through the A*MIDEX project (n°ANR-11-IDEX-0001-02).

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