Stimulation of the Serotonin Receptor 7 Restores Brain Histone H3 Acetylation and MeCP2 Corepressor Protein Levels in a Female Mouse Model of Rett Syndrome

J Neuropathol Exp Neurol. 2021 Feb 22;80(3):265-273. doi: 10.1093/jnen/nlaa158.

Abstract

Rett syndrome (RTT) is a rare neurological disorder caused by mutations in the X-linked MECP2 gene, characterized by severe behavioral and physiological impairments for which no cure is available. The stimulation of serotonin receptor 7 (5-HT7R) with its selective agonist LP-211 (0.25 mg/kg/day for 7 days) was proved to rescue neurobehavioral alterations in a mouse model of RTT. In the present study, we aimed at gaining insight into the mechanisms underpinning the efficacy of 5-HT7R pharmacological stimulation by investigating its epigenetic outcomes in the brain of RTT female mice bearing a truncating MeCP2 mutation. Treatment with LP-211 normalized the reduced histone H3 acetylation and HDAC3/NCoR levels, and increased HDAC1/Sin3a expression in RTT mouse cortex. Repeated 5-HT7R stimulation also appeared to strengthen the association between NCoR and MeCP2 in the same brain region. A different profile was found in RTT hippocampus, where LP-211 rescued H3 hyperacetylation and increased HDAC3 levels. Overall, the present data highlight a new scenario on the relationship between histone acetylation and serotoninergic pathways. 5-HT7R is confirmed as a pivotal therapeutic target for the recovery of neuronal function supporting the translational value of this promising pharmacological approach for RTT.

Keywords: Brain plasticity; Corepressor complexes; Histone acetylation; MeCP2; Rett syndrome; Serotonin receptor 7.

Publication types

  • Research Support, Non-U.S. Gov't

MeSH terms

  • Acetylation
  • Animals
  • Brain / drug effects
  • Brain / metabolism*
  • Disease Models, Animal*
  • Female
  • Histones / genetics
  • Histones / metabolism*
  • Methyl-CpG-Binding Protein 2 / genetics
  • Methyl-CpG-Binding Protein 2 / metabolism*
  • Mice
  • Mice, Inbred C57BL
  • Mice, Transgenic
  • Piperazines / pharmacology
  • Piperazines / therapeutic use
  • Receptors, Serotonin / metabolism*
  • Rett Syndrome / drug therapy
  • Rett Syndrome / genetics
  • Rett Syndrome / metabolism*
  • Serotonin Receptor Agonists / pharmacology
  • Serotonin Receptor Agonists / therapeutic use

Substances

  • Histones
  • Mecp2 protein, mouse
  • Methyl-CpG-Binding Protein 2
  • N-(4-cyanophenylmethyl)-4-(2-diphenyl)-1-piperazinehexanamide
  • Piperazines
  • Receptors, Serotonin
  • Serotonin Receptor Agonists
  • serotonin 7 receptor