1 Ministry of Education Key Laboratory of Contemporary Anthropology, Department of Anthropology and Human Genetics, School of Life Sciences, Fudan University, Shanghai, 200438, China.
2 Guizhou Municipal Institute of Cultural Relics and Archaeology, Guiyang, 5500044, China. Electronic address: 1513051739@qq.com.
3 Department of Integrative Anatomical Sciences, Keck School of Medicine, University of Southern California, Los Angeles, CA 90033, USA; Evolutionary Studies Institute, University of the Witwatersrand, WITS 2050, Johannesburg, South Africa. Electronic address: kristian.carlson@usc.edu.
4 Guizhou Municipal Institute of Cultural Relics and Archaeology, Guiyang, 5500044, China.
5 State Key Laboratory of Genetic Engineering, Collaborative Innovation Center for Genetics and Development, School of Life Sciences, Fudan University, Shanghai, 200438, China; Human Phoneme Institute, Fudan University, Shanghai, 200438, China.
6 Key Laboratory of Vertebrate Evolution and Human Origins, Institute of Vertebrate Palaeontology and Palaeoanthropology, Chinese Academy of Sciences, Beijing 100044, China; CAS Center for Excellence in Life and Paleoenvironment, Beijing 100044, China.