1 MEMO Research, Division of Molecular and Clinical Medicine, University of Dundee, Dundee DD1 9SY, UK. Electronic address: t.m.macdonald@dundee.ac.uk.
2 Centre for Genomic and Experimental Medicine, University of Edinburgh, Edinburgh, UK.
3 The Robertson Centre for Biostatistics, University of Glasgow, Glasgow, UK.
4 MEMO Research, Division of Molecular and Clinical Medicine, University of Dundee, Dundee DD1 9SY, UK.