1 Naturalis Biodiversity Center, Postbus 9517, 2300 RA Leiden, The Netherlands.. g.sangster@planet.nl.
2 Animal Ecology, Department of Ecology and Genetics, Evolutionary Biology Centre, Uppsala University, Norbyvgen 18 D, SE-752 36 Uppsala, Sweden. user@example.com.
3 34, avenue Antoine de Saint-Exupry, 17 000 La Rochelle, France. . user@example.com.
4 Department of Biological and Environmental Sciences, University of Gteborg, Box 463, SE-413 90 Gothenburg, Sweden. Gothenburg Global Biodiversity Centre, Box 461, SE-405 30 Gothenburg, Sweden.. user@example.com.