1 Department of Respiratory Medicine and Clinical Immunology, Osaka University Graduate School of Medicine, Osaka, Japan.
2 Department of Otorhinolaryngology-Head and Neck Surgery, Osaka University Graduate School of Medicine, Osaka, Japan.
3 Department of Pathology, Osaka University Graduate School of Medicine, Osaka, Japan.
4 Department of Immunopathology, WPI, Immunology Frontier Research Center (iFReC) Osaka University, Osaka, Japan.
5 Integrated Frontier Research for Medical Science Division, Institute for Open and Transdisciplinary Research Initiatives (OTRI) Osaka University, Osaka, Japan.
6 Center for Infectious Disease for Education and Research (CiDER), Osaka University, Osaka, Japan.