1 Manchester Royal Eye Hospital and Manchester Academic Health Sciences Centre, Manchester University NHS Foundation Trust Manchester, Manchester, M13 9WL, UK. robert.harper@mft.nhs.uk.
2 Division of Pharmacy and Optometry, School of Health Sciences, Faculty of Biology, Medicine and Health, University of Manchester, Manchester, M13 9PL, UK. robert.harper@mft.nhs.uk.
3 Conclusio, Ty Derw Lime Tree Court, Cardiff Gate Business Park, Cardiff, CF23 8AB, UK.
4 Manchester Royal Eye Hospital and Manchester Academic Health Sciences Centre, Manchester University NHS Foundation Trust Manchester, Manchester, M13 9WL, UK.
5 Division of Pharmacy and Optometry, School of Health Sciences, Faculty of Biology, Medicine and Health, University of Manchester, Manchester, M13 9PL, UK.
6 Division of Evolution and Genomic Sciences, School of Health Sciences, Faculty of Biology, Medicines and Health, University of Manchester, Manchester, M13 9PL, UK.