A single-nucleus and spatial transcriptomic atlas of the COVID-19 liver reveals topological, functional, and regenerative organ disruption in patients

Genome Biol. 2025 Mar 14;26(1):56. doi: 10.1186/s13059-025-03499-5.

Abstract

Background: The molecular underpinnings of organ dysfunction in severe COVID-19 and its potential long-term sequelae are under intense investigation. To shed light on these in the context of liver function, we perform single-nucleus RNA-seq and spatial transcriptomic profiling of livers from 17 COVID-19 decedents.

Results: We identify hepatocytes positive for SARS-CoV-2 RNA with an expression phenotype resembling infected lung epithelial cells, and a central role in a pro-fibrotic TGFβ signaling cell-cell communications network. Integrated analysis and comparisons with healthy controls reveal extensive changes in the cellular composition and expression states in COVID-19 liver, providing the underpinning of hepatocellular injury, ductular reaction, pathologic vascular expansion, and fibrogenesis characteristic of COVID-19 cholangiopathy. We also observe Kupffer cell proliferation and erythrocyte progenitors for the first time in a human liver single-cell atlas. Despite the absence of a clinical acute liver injury phenotype, endothelial cell composition is dramatically impacted in COVID-19, concomitantly with extensive alterations and profibrogenic activation of reactive cholangiocytes and mesenchymal cells.

Conclusions: Our atlas provides novel insights into liver physiology and pathology in COVID-19 and forms a foundational resource for its investigation and understanding.

Keywords: COVID-19; Liver; SARS-CoV-2; Single-cell sequencing; Single-nucleus sequencing; Spatial transcriptomics.

MeSH terms

  • Aged
  • COVID-19* / genetics
  • COVID-19* / metabolism
  • COVID-19* / pathology
  • COVID-19* / virology
  • Female
  • Gene Expression Profiling
  • Hepatocytes / metabolism
  • Hepatocytes / pathology
  • Hepatocytes / virology
  • Humans
  • Liver* / metabolism
  • Liver* / pathology
  • Liver* / virology
  • Male
  • Middle Aged
  • SARS-CoV-2 / genetics
  • Single-Cell Analysis
  • Transcriptome*