1 Sir William Dunn School of Pathology, University of Oxford, Oxford, UK.
2 Oxford Protein Informatics Group, Department of Statistics, University of Oxford, Oxford, UK.
3 Department of Mathematics and Statistics, Boston University, Boston, Massachusetts, USA.
4 Kirby Institute, University of New South Wales, Sydney, New South Wales, Australia.
5 MRC Translate Immune Discovery Unit, MRC Weatherall Institute of Molecular Medicine, University of Oxford, Oxford, UK.
6 Department of Mathematics and Department of Biology, Northeastern University, Boston, Massachusetts, USA.
7 Wellcome Centre for Human Genetics, Nuffield Department of Medicine, University of Oxford, Oxford, UK.
8 Chinese Academy of Medical Science Oxford Institute, University of Oxford, Oxford, UK.
9 Division of Emerging Infectious Disease, Research Department, Faculty of Medicine Siriraj Hospital, Mahidol University, Bangkoknoi, Bangkok, Thailand.
10 James & Lillian Martin Centre, Sir William Dunn School of Pathology, University of Oxford, Oxford, UK.
11 Oxford University Hospitals NHS Foundation Trust, Oxford, Oxford, UK.
12 Oxford Protein Informatics Group, Department of Statistics, University of Oxford, Oxford, UK. deane@stats.ox.ac.uk.
13 Department of Mathematics and Statistics, Boston University, Boston, Massachusetts, USA. isaacsas@bu.edu.
14 Sir William Dunn School of Pathology, University of Oxford, Oxford, UK. omer.dushek@path.ox.ac.uk.