Profiling of human IL-22+ T cell clones from patients affected with Schistosoma mansoni: Insights into macrophage regulation and liver fibrosis

PLoS Negl Trop Dis. 2025 May 30;19(5):e0013132. doi: 10.1371/journal.pntd.0013132. eCollection 2025 May.

Abstract

Tissue damage in Schistosoma mansoni infection results from a granulomatous, T cell-mediated response to parasite eggs, leading to liver fibrosis and portal hypertension. This immune response, initially Th1-dominated, progressively shifts toward a Th2 profile, contributing to hepatic stellate cell (HSC) activation and fibrosis. However, the precise regulatory mechanisms remain unclear. In this study, we analyzed T cell responses to soluble egg antigens (SEA) in 121 T cell clones (Tcc) from S. mansoni-infected patients. All clones produced high levels of IL-13 upon anti-CD3 stimulation; a minority secreted IFN-γ (n = 33) or IL-10 (n = 38). Notably, 51 clones co-produced IL-22 and IL-13. To investigate IL-22's role, we examined IL-22 receptor (IL-22R) expression on human M0 and M2 macrophages. Both subsets expressed IL-22R, and its engagement triggered phosphorylation of p38, STAT3, and STAT5. IL-22 also downregulated IL-13-induced M2 markers (CD163, CD200R). Furthermore, IL-22 treatment of HSCs inhibited IL-13-driven collagen I/III production and cell proliferation. These results suggest that IL-22-producing T cells modulate Th2 macrophage polarization and directly suppress fibrogenesis in HSCs. IL-22 may thus act as a regulatory cytokine counteracting liver fibrosis during schistosomiasis.

MeSH terms

  • Adult
  • Animals
  • Antigens, Helminth / immunology
  • Female
  • Hepatic Stellate Cells / immunology
  • Humans
  • Interleukin-13 / metabolism
  • Interleukin-22
  • Interleukins* / immunology
  • Interleukins* / metabolism
  • Liver Cirrhosis* / immunology
  • Liver Cirrhosis* / parasitology
  • Macrophages* / immunology
  • Male
  • Middle Aged
  • Schistosoma mansoni* / immunology
  • Schistosomiasis mansoni* / immunology
  • Schistosomiasis mansoni* / pathology
  • T-Lymphocytes* / immunology

Substances

  • Interleukins
  • Interleukin-22
  • Interleukin-13
  • Antigens, Helminth