Systematic identification of single transcription factor perturbations that drive cellular and tissue rejuvenation

Proc Natl Acad Sci U S A. 2026 Jan 13;123(2):e2515183123. doi: 10.1073/pnas.2515183123. Epub 2026 Jan 9.

Abstract

Cellular rejuvenation through transcriptional reprogramming is an exciting approach to counter aging. Using a fibroblast-based model of human cell aging and Perturb-seq screening, we developed a systematic approach to identify single transcription factor (TF) perturbations that promote rejuvenation without dedifferentiation. Overexpressing E2F3 or EZH2, and repressing STAT3 or ZFX, reversed cellular hallmarks of aging-increasing proliferation, proteostasis, and mitochondrial activity, while decreasing senescence. EZH2 overexpression in vivo rejuvenated livers in aged mice, reversing aging-associated gene expression profiles, decreasing steatosis and fibrosis, and improving glucose tolerance. Mechanistically, single TF perturbations led to convergent downstream transcriptional programs conserved in different aging and rejuvenation models. These results suggest a shared set of molecular requirements for cellular and tissue rejuvenation across species.

Keywords: Perturb-seq screening; liver aging; rejuvenation; replicative aging.

MeSH terms

  • Aging / genetics
  • Animals
  • Cell Proliferation
  • Cellular Senescence / genetics
  • Enhancer of Zeste Homolog 2 Protein / genetics
  • Enhancer of Zeste Homolog 2 Protein / metabolism
  • Fibroblasts / metabolism
  • Humans
  • Liver / metabolism
  • Mice
  • Rejuvenation* / physiology
  • STAT3 Transcription Factor / genetics
  • STAT3 Transcription Factor / metabolism
  • Transcription Factors* / genetics
  • Transcription Factors* / metabolism

Substances

  • Enhancer of Zeste Homolog 2 Protein
  • Transcription Factors
  • STAT3 Transcription Factor
  • EZH2 protein, human