1 Laboratory of Microbiology, Department of Disease Control, Faculty of Veterinary Medicine, Hokkaido University, Kita 18, Nishi 9, Kita-Ku, Sapporo, Hokkaido, 060-0818, Japan.
2 Laboratory of Microbiology, Department of Disease Control, Faculty of Veterinary Medicine, Hokkaido University, Kita 18, Nishi 9, Kita-Ku, Sapporo, Hokkaido, 060-0818, Japan. hiono@vetmed.hokudai.ac.jp.
3 One Health Research Center, Hokkaido University, Sapporo, Hokkaido, 060-0818, Japan. hiono@vetmed.hokudai.ac.jp.
4 Hokkaido University Institute for Vaccine Research and Development (HU-IVReD), Hokkaido University, Sapporo, Hokkaido, 001-0021, Japan. hiono@vetmed.hokudai.ac.jp.
5 International Collaboration Unit, International Institute for Zoonosis Control, Hokkaido University, Sapporo, Hokkaido, 001-0020, Japan. hiono@vetmed.hokudai.ac.jp.
6 Area for Molecular Function, Division of Material Science, Graduate School of Science and Engineering, Saitama University, 225 Shimo-Ohkubo, Sakura-Ku, Saitama, Saitama, 338-8570, Japan.
7 International Collaboration Unit, International Institute for Zoonosis Control, Hokkaido University, Sapporo, Hokkaido, 001-0020, Japan.
8 Division of Global Epidemiology, International Institute for Zoonosis Control, Hokkaido University, Kita 20, Nishi 10, Kita-Ku, Sapporo, Hokkaido, 001-0020, Japan.
9 Medical Innovation Research Unit (MiU), Advanced Institute of Innovative Technology (AIIT), Saitama University, Saitama, Saitama, 338-8570, Japan.
10 Area for Health Science, Strategic Research Center, Saitama University, Saitama, Saitama, 338-8570, Japan.
11 One Health Research Center, Hokkaido University, Sapporo, Hokkaido, 060-0818, Japan.
12 Hokkaido University Institute for Vaccine Research and Development (HU-IVReD), Hokkaido University, Sapporo, Hokkaido, 001-0021, Japan.