1 Department of Bioscience, School of Biological and Environmental Sciences, Kwansei Gakuin University, 1 Gakuen-Uegahara, Sanda, 669-1330, Hyogo, Japan.
2 Biotechnology Laboratory, Electric Power Development Co., Ltd, Kitakyushu, Japan.
3 Division of Biotechnology and Life Science, Institute of Engineering, Tokyo University of Agriculture and Technology, 2-24-16, Koganei, 184-8588, Tokyo, Japan.
4 Department of Bioscience, School of Biological and Environmental Sciences, Kwansei Gakuin University, 1 Gakuen-Uegahara, Sanda, 669-1330, Hyogo, Japan. gshimakawa@panda.kobe-u.ac.jp.
5 Graduate School of Agricultural Science, Kobe University, 1-1 Rokkodai, Nada, Kobe, 657-8501, Japan. gshimakawa@panda.kobe-u.ac.jp.
6 Department of Bioscience, School of Biological and Environmental Sciences, Kwansei Gakuin University, 1 Gakuen-Uegahara, Sanda, 669-1330, Hyogo, Japan. yusuke@kwansei.ac.jp.