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Walls are thin 1 (WAT1), an Arabidopsis homolog of Medicago truncatula NODULIN21, is a tonoplast-localized protein required for secondary wall formation in fibers.
Ranocha P, Denancé N, Vanholme R, Freydier A, Martinez Y, Hoffmann L, Köhler L, Pouzet C, Renou JP, Sundberg B, Boerjan W, Goffner D. Ranocha P, et al. Among authors: kohler l. Plant J. 2010 Aug;63(3):469-83. doi: 10.1111/j.1365-313X.2010.04256.x. Epub 2010 May 20. Plant J. 2010. PMID: 20497379 Free article.