1 QIMR Berghofer Medical Research Institute, Brisbane, Queensland, Australia.
2 QIMR Berghofer Medical Research Institute, Brisbane, Queensland, Australia; Genetics and Immunology Group, University of the Highlands and Islands, Inverness, United Kingdom.
3 QIMR Berghofer Medical Research Institute, Brisbane, Queensland, Australia. Electronic address: Nicholas.Hayward@qimrberghofer.edu.au.