1 Laboratory for Circuit and Behavioral Physiology, RIKEN, Japan. Electronic address: thomas.mchugh@riken.jp.
2 Institute of Neuroscience, Center for Excellence in Brain Science and Intelligent Technology, Chinese Academy of Sciences, Shanghai, China; Shanghai Center for Brain Science and Brain-Inspired Technology, Center for Brain Science, Wakoshi, Saitama, Japan. Electronic address: mpoo@ion.ac.cn.